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The role that plastoquinone plays in photosynthesis, more specifically in the light-dependent reactions of photosynthesis, is that of a mobile electron carrier through the membrane of the thylakoid. [2] Plastoquinone is reduced when it accepts two electrons from photosystem II and two hydrogen cations (H +) from the stroma of the chloroplast ...
Photosystem II (or water-plastoquinone oxidoreductase) is the first protein complex in the energy-dependent reactions of oxygenic photosynthesis. It is located in the thylakoid membrane of plants , algae , and cyanobacteria .
To do this, it must release the absorbed energy. This can happen in various ways. The extra energy can be converted into molecular motion and lost as heat, or re-emitted by the electron as light (fluorescence). The energy, but not the electron itself, may be passed onto another molecule; this is called resonance energy transfer.
The thylakoid membranes of higher plants are composed primarily of phospholipids [5] and galactolipids that are asymmetrically arranged along and across the membranes. [6] Thylakoid membranes are richer in galactolipids rather than phospholipids; also they predominantly consist of hexagonal phase II forming monogalacotosyl diglyceride lipid.
The cytochrome b 6 f complex is a dimer, with each monomer composed of eight subunits. [3] These consist of four large subunits: a 32 kDa cytochrome f with a c-type cytochrome, a 25 kDa cytochrome b 6 with a low- and high-potential heme group, a 19 kDa Rieske iron-sulfur protein containing a [2Fe-2S] cluster, and a 17 kDa subunit IV; along with four small subunits (3-4 kDa): PetG, PetL, PetM ...
When the photon has been absorbed, the resulting high-energy electron is transferred to a nearby pheophytin molecule. This is above and to the right of the pair on the diagram and is coloured grey. The electron travels from the pheophytin molecule through two plastoquinone molecules, the first tightly bound, the second loosely bound.
The electrons then pass through Cyt b 6 and Cyt f to plastocyanin, using energy from photosystem I to pump hydrogen ions (H +) into the thylakoid space. This creates a H + gradient, making H + ions flow back into the stroma of the chloroplast, providing the energy for the (re)generation of ATP.
The pigments which absorb light at the highest energy level are found furthest from the reaction center. On the other hand, the pigments with the lowest energy level are more closely associated with the reaction center. Energy will be efficiently transferred from the outer part of the antenna complex to the inner part.