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These organisms perform photosynthesis through organelles called chloroplasts and are believed to have originated about 2 billion years ago. [1] Comparing the genes of chloroplast and cyanobacteria strongly suggests that chloroplasts evolved as a result of endosymbiosis with cyanobacteria that gradually lost the genes required to be free-living.
Possible cladogram of chloroplast evolution [2] [3] Circles represent endosymbiotic events. For clarity, dinophyte tertiary endosymbioses and many nonphotosynthetic lineages have been omitted. a It is now established that Chromalveolata is paraphyletic to Rhizaria .
Chloroplasts, containing thylakoids, visible in the cells of Ptychostomum capillare, a type of moss. A chloroplast (/ ˈ k l ɔːr ə ˌ p l æ s t,-p l ɑː s t /) [1] [2] is a type of organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells.
The evolution of photosynthesis refers to the origin and subsequent evolution of photosynthesis, the process by which light energy is used to assemble sugars from carbon dioxide and a hydrogen and electron source such as water. It is believed that the pigments used for photosynthesis initially were used for protection from the harmful effects ...
The Russian botanist Konstantin Mereschkowski first outlined the theory of symbiogenesis (from Greek: σύν syn "together", βίος bios "life", and γένεσις genesis "origin, birth") in his 1905 work, The nature and origins of chromatophores in the plant kingdom, and then elaborated it in his 1910 The Theory of Two Plasms as the Basis ...
Comparatively, in the laboratory, most cultured cells—which are large compared to normal plant cells—produce very long and abundant stromules that extend to the cell periphery. In 2014, evidence was found of the possible loss of plastid genome in Rafflesia lagascae , a non-photosynthetic parasitic flowering plant, and in Polytomella , a ...
Despite this, chloroplasts can be found in an extremely wide set of organisms, some not even directly related to each other—a consequence of many secondary and even tertiary endosymbiotic events. The chloroplasts of glaucophytes have a peptidoglycan layer, evidence suggesting their endosymbiotic origin from cyanobacteria .
Each of the envelope membranes is a lipid bilayer that is between 6 and 8 nm thick. The lipid composition of the outer membrane has been found to be 48% phospholipids, 46% galactolipids and 7% sulfolipids, while the inner membrane has been found to contain 16% phospholipids, 79% galactolipids and 5% sulfolipids in spinach chloroplasts.