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In this spider diagram, the position of the book lungs is labelled 1. Spider book lungs (cross section) Internal anatomy of a female spider, book lungs shown in pink A book lung is a type of respiration organ used for atmospheric gas-exchange that is present in many arachnids, such as scorpions and spiders.
Although all insects have spiracles, only some arachnids have them. Some spiders such as orb weavers and wolf spiders have spiracles. Ancestrally, spiders have book lungs, not trachea. However, some spiders evolved a tracheal system independently of the tracheal system in insects, which includes independent evolution of the spiracles as well ...
Insects have spiracles on their exoskeletons to allow air to enter the trachea. [1] [page needed] In insects, the tracheal tubes primarily deliver oxygen directly into the insects' tissues. The spiracles can be opened and closed in an efficient manner to reduce water loss. This is done by contracting closer muscles surrounding the spiracle.
There are characteristics that are particularly important for the terrestrial lifestyle of arachnids, such as internal respiratory surfaces in the form of tracheae, or modification of the book gill into a book lung, an internal series of vascular lamellae used for gas exchange with the air. [21]
Crustacea usually have gills that are modified appendages. Many arachnids have book lungs. [63] Tracheae, systems of branching tunnels that run from the openings in the body walls, deliver oxygen directly to individual cells in many insects, myriapods and arachnids. [64]
These depend on individual sub-groups' environments. Modern terrestrial chelicerates generally have both book lungs, which deliver oxygen and remove waste gases via the blood, and tracheae, which do the same without using the blood as a transport system. [23] The living horseshoe crabs are aquatic and have book gills that
Spider book lungs (cross section) Spiders have developed several different respiratory anatomies, based either on book lungs or on tracheae. Mesothele and mygalomorph spiders have two pairs of book lungs filled with haemolymph, where openings on the ventral surface of the abdomen allow air to enter and oxygen to diffuse in and carbon dioxide to ...
In animals like whip scorpions and whip spiders the first two 'sternites' bearing the book lungs may actually be highly modified opisthosomal limbs. Segmentation of the opisthosoma in adult spiders is not visible, but embryo spiders typically have 13 segments, the posterior segments being called the presegmental zone.