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The butterflies form the clade Rhopalocera, which is composed of three superfamilies: Hedyloidea (the moth butterfly family Hedylidae), the Hesperioidea (the skipper family Hesperiidae), and the Papilionoidea (the true butterfly families Papilionidae, Pieridae, Nymphalidae, Lycaenidae, and Riodinidae). All of these families are monophyletic.
Butterflies evolved from moths, so while the butterflies are monophyletic (forming a single clade), the moths are not. The oldest known butterfly is Protocoeliades kristenseni from the Palaeocene aged Fur Formation of Denmark, approximately 55 million years old, which belongs to the family Hesperiidae (skippers). [ 9 ]
Modern-type butterflies and moths appear. Extinction of Gastornis. Basilosaurus, one of the first of the giant whales, appeared in the fossil record. 38 Ma Earliest bears. 37 Ma First nimravid ("false saber-toothed cats") carnivores — these species are unrelated to modern-type felines. First alligators and ruminants. 35 Ma
Plants evolved chemical defenses against this herbivory and the insects in turn evolved mechanisms to deal with plant toxins. [5] These toxins limit the diet breadth of herbivores, and evolving mechanisms to nonetheless continue herbivory is an important part of maintaining diet breadth in insects, and so in their evolutionary history as a whole.
Lepidoptera (/ ˌ l ɛ p ɪ ˈ d ɒ p t ər ə / LEP-ih-DOP-tər-ə) or lepidopterans is an order of winged insects which includes butterflies and moths.About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organisms, [1] [2] making it the second largest insect order (behind Coleoptera) with 126 families [3] and 46 superfamilies ...
A = Anapsid, B = Synapsid, C = Diapsid. It was traditionally assumed that first reptiles were anapsids, having a solid skull with holes only for the nose, eyes, spinal cord, etc.; [10] the discoveries of synapsid-like openings in the skull roof of the skulls of several members of Parareptilia, including lanthanosuchoids, millerettids, bolosaurids, some nycteroleterids, some procolophonoids and ...
The Rhopalocera, comprising the Papilionoidea (butterflies), Hesperioidea (skippers), and the Hedyloidea (moth-butterflies), are the most recently evolved. [9] There is quite a good fossil record for this group, with the oldest skipper dating from 56 million years ago .
Owl butterflies are very large, 65–200 mm (2.6–7.9 in), and fly only a few meters at a time, so avian predators have little difficulty in following them to their settling place. However, the butterflies preferentially fly in dusk, when few avian predators are around. [ 1 ]