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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
In the stems of some Asterales dicots, there may be phloem located inwardly from the xylem as well. Between the xylem and phloem is a meristem called the vascular cambium. This tissue divides off cells that will become additional xylem and phloem. This growth increases the girth of the plant, rather than its length.
Bast fibre (also called phloem fibre or skin fibre) is plant fibre collected from the phloem (the "inner bark", sometimes called "skin") or bast surrounding the stem of certain dicotyledonous plants.
Phloem and xylem are closely associated with one another and are typically located immediately adjacent to each other in the plant. The combination of one xylem and one phloem strand adjacent to each other is known as a vascular bundle . [ 14 ]
Phloem was introduced by Carl Nägeli in 1858 after the discovery of sieve elements. Since then, multiple studies have been conducted on how sieve elements function in phloem in terms of working as a transport mechanism. [2] An example of analysis of phloem through sieve elements was conducted in the study of Arabidopsis leaves.
Phloem is a complex tissue, consisting of two main cell types, the sieve tubes and the intimately associated companion cells, together with parenchyma cells, phloem fibres and sclereids. [ 19 ] : 171 Sieve tubes are joined end-to-end with perforated end-plates between known as sieve plates , which allow transport of photosynthate between the ...
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The fascicular and interfascicular cambia thus join up to form a ring (in three dimensions, a tube) which separates the primary xylem and primary phloem, the cambium ring. The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart.