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The Russian tortoise (Testudo horsfieldii), also commonly known as the Afghan tortoise, the Central Asian tortoise, the four-clawed tortoise, the four-toed tortoise, Horsfield's tortoise, the Russian steppe tortoise, the Soviet Tortoise, and the steppe tortoise, [3] [4] is a threatened species of tortoise in the family Testudinidae.
The following extant species in the following subgenera are placed here: Genus Testudo. Subgenus Agrionemys. Russian tortoise or Horsfield's tortoise, T. horsfieldii [1] Subspecies: Central Asian tortoise, T. horsfieldii horsfieldii; Fergana Valley steppe tortoise, T. horsfieldii bogdanovi; Kazakhstan steppe tortoise, T. horsfieldii kazakhstanica
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Although some factors indicate this might be correct, [6] the data at hand are not unequivocally in support and the relationships between Hermann's and the Russian tortoise among each other and to the other species placed in Testudo are not robustly determined. Hence, it seems doubtful that the new genus will be accepted for now.
Tortoises are the longest-living land animals in the world, although the longest-living species of tortoise is a matter of debate. Galápagos tortoises are noted to live over 150 years, but an Aldabra giant tortoise named Adwaita may have lived an estimated 255 years. In general, most tortoise species can live 80–150 years.
The other four species are Hermann's tortoise (T. hermanni), the Egyptian tortoise (T. kleinmanni), the marginated tortoise (T. marginata), and the Russian tortoise (A. horsfieldii). The Greek tortoise is a very long-lived animal, achieving a lifespan upwards of 125 years, with some unverified reports up to 200 years. [4]
Red-footed tortoises have many common names: red-leg, red-legged, or red-foot tortoise (often without the hyphen) and the savanna tortoise, as well as local names, such as carumbe or karumbe, which means 'slow moving' (Brazil, Paraguay), wayapopi or morrocoy (Venezuela, Colombia), and variations of jabuti such as japuta and jabuti-piranga (Brazil, Argentina). [5]
The ancient tortoises had primitive jaw muscles, unlike today's tortoises, which also display the os transiliens bone, and would have been herbivorous. While Stylemys species did exhibit the same neck structure as modern tortoises, the forelimbs were unsuitable for burrowing, setting them apart from modern genera.