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The term monophyly, or monophyletic, derives from the two Ancient Greek words μόνος (mónos), meaning "alone, only, unique", and φῦλον (phûlon), meaning "genus, species", [4] [5] and refers to the fact that a monophyletic group includes organisms (e.g., genera, species) consisting of all the descendants of a unique common ancestor.
In biological phylogenetics, a clade (from Ancient Greek κλάδος (kládos) 'branch'), also known as a monophyletic group or natural group, [1] is a grouping of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. [2]
Willi Hennig 1972 Peter Chalmers Mitchell in 1920 Robert John Tillyard. The original methods used in cladistic analysis and the school of taxonomy derived from the work of the German entomologist Willi Hennig, who referred to it as phylogenetic systematics (also the title of his 1966 book); but the terms "cladistics" and "clade" were popularized by other researchers.
While the butterflies form a monophyletic group, the moths, which comprise the rest of the Lepidoptera, do not. Many attempts have been made to group the superfamilies of the Lepidoptera into natural groups, most of which fail because one of the two groups is not monophyletic: Microlepidoptera and Macrolepidoptera, Heterocera and Rhopalocera, Jugatae and Frenatae, Monotrysia and Ditrysia.
They held that only monophyletic groups should be accepted as formal ranks in a classification and that – while this approach had been impractical previously (necessitating "literally dozens of eukaryotic 'kingdoms '") – it had now become possible to divide the eukaryotes into "just a few major groups that are probably all monophyletic". [42]
Flower of Liriodendron tulipifera, a Mesangiosperm. Mesangiospermae is a clade that contains the majority of flowering plants (angiosperms). Mesangiosperms are therefore known as the core angiosperms, in contrast to the three orders of earlier-diverging species known as the basal angiosperms: Nymphaeales (including water lilies), Austrobaileyales (including star anise), and Amborellales.
The ranks of higher taxa, especially intermediate ranks, are prone to revision as new information about relationships is discovered. For example, the flowering plants have been downgraded from a division (Magnoliophyta) to a subclass (Magnoliidae), and the superorder has become the rank that distinguishes the major groups of flowering plants. [23]
"Tricolpate" is a synonym for the "Eudicot" monophyletic group, the "true dicotyledons" (which are distinguished from all other flowering plants by their tricolpate pollen structure). The number of pollen grain furrows or pores helps classify the flowering plants, with eudicots having three colpi (tricolpate), and other groups having one sulcus ...