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Agnatha (/ ˈ æ ɡ n ə θ ə, æ ɡ ˈ n eɪ θ ə /; [3] from Ancient Greek ἀ-(a-) 'without' and γνάθος (gnáthos) 'jaws') is a paraphyletic infraphylum [4] of non-gnathostome vertebrates, or jawless fish, in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and extinct (conodonts, anaspids, and ostracoderms, among others).
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Cyclostomi, often referred to as Cyclostomata / s ɪ k l oʊ ˈ s t ɒ m ə t ə /, is a group of vertebrates that comprises the living jawless fishes: the lampreys and hagfishes.Both groups have jawless mouths with horny epidermal structures that function as teeth called ceratodontes, and branchial arches that are internally positioned instead of external as in the related jawed fishes. [1]
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Together, these two taxa of jawless vertebrates are the sister-group of gnathostomes. Several synapomorphies support this hypothesis, such as the presence of: sclerotic ossicles, paired pectoral fins, a dermal skeleton with three layers (a basal layer of isopedin, a middle layer of spongy bone, and a superficial layer of dentin), and ...
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Recent molecular and morphological phylogenetic studies place lampreys and hagfish in the infraphylum Agnatha or Agnathostomata (both meaning without jaws). The other vertebrate infraphylum is Gnathostomata (jawed mouths) and includes the classes Chondrichthyes (sharks), Osteichthyes (bony fishes), Amphibia, Reptilia, Aves, and Mammalia.
Gnathostomata is traditionally an infraphylum, broken into three top-level groupings: Chondrichthyes, or the cartilaginous fish; Placodermi, an extinct grade of armored fish; and Teleostomi, which includes the familiar classes of bony fish, birds, mammals, reptiles, and amphibians. Some classification systems have used the term Amphirhina.