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The endodermis is the boundary between the cortex and the stele. In many seedless plants, such as ferns, the endodermis is a distinct layer of cells immediately outside the vascular cylinder (stele) in roots and shoots. In most seed plants, especially woody types, the endodermis is present in roots but not in stems.
The earliest vascular plants had stems with a central core of vascular tissue. [3] [4] This consisted of a cylindrical strand of xylem, surrounded by a region of phloem. Around the vascular tissue there might have been an endodermis that regulated the flow of water into and out of the vascular system. Such an arrangement is termed a protostele.
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. [1] The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. [2]
The following description is for early events in lateral root formation of the model organism Arabidopsis thaliana: Lateral root formation is initiated in pericycle (located between the endodermis and vascular tissue) of the root system, and begins with a process referred to as priming. In this stage, you have rhythmic bouts of gene expression ...
In dicot root, the vascular cambium is completely secondary in origin, and it originates from a portion of pericycle tissue. [citation needed] The pericycle regulates the formation of lateral roots by rapidly dividing near the xylem elements of the root. [2] It has been known to often be confused with other parts of the plant.
These bundles include both xylem and phloem, as well as supporting and protective cells. In stems and roots, the xylem typically lies closer to the interior of the stem with phloem towards the exterior of the stem. In the stems of some Asterales dicots, there may be phloem located inwardly from the xylem as well.
The Casparian strip is mainly located in the endodermis of the root, [26] but some plants also have the Casparian strip in the outer cortex on the outer side of the root cortex, stem or leaf. [27] For example, the conifers of Pinus bungeana and the stems of Pelargonium have the Casparian strip, which may be related to preventing water ...
Roots are specialized for the uptake of water, nutrients (including ions for proper function). [12] Similar to the endodermis, the exodermis contains very compact cells and is surrounded by a Casparian band, two features which are used to restrict the flow of water to a symplastic fashion (through the cytoplasm) rather than apoplastic fashion which (through the cell wall) flow through passages ...