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Lignin fills the spaces in the cell wall between cellulose, hemicellulose, and pectin components, especially in vascular and support tissues: xylem tracheids, vessel elements and sclereid cells. [citation needed] Lignin plays a crucial part in conducting water and aqueous nutrients in plant stems.
The inclusion of lignin makes the secondary cell wall less flexible and less permeable to water than the primary cell wall. [4] In addition to making the walls more resistant to degradation, the hydrophobic nature of lignin within these tissues is essential for containing water within the vascular tissues that carry it throughout the plant.
The basic function of the xylem is to transport water upward from the roots to parts of the plants such as stems and leaves, but it also transports nutrients. [1] [2] The word xylem is derived from the Ancient Greek word, ξύλον (xylon), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. [3]
All the vascular tissues within a particular plant together constitute the vascular tissue system of that plant. The cells in vascular tissue are typically long and slender. Since the xylem and phloem function in the conduction of water, minerals, and nutrients throughout the plant, it is not surprising that their form should be similar to pipes.
Sclerenchyma is the tissue which makes the plant hard and stiff. Sclerenchyma is the supporting tissue in plants. Two types of sclerenchyma cells exist: fibers cellular and sclereids. Their cell walls consist of cellulose, hemicellulose, and lignin. Sclerenchyma cells are the principal supporting cells in plant tissues that have ceased elongation.
The xylem consists of vessels in flowering plants and of tracheids in other vascular plants. Xylem cells are dead, hard-walled hollow cells arranged to form files of tubes that function in water transport. A tracheid cell wall usually contains the polymer lignin. The phloem, on the other hand, consists of living cells called sieve-tube members ...
The rigid structure of the lignin protein gives a sturdy structure to the tube, and even provides some structure and support for the plant. Xylem mainly functions to transport water from the roots to the rest of the plant, however it also transports some nutrients, such as amino acids, small proteins, ions, and some other vital nutrients. [8]
In the first 140–150 million years of vascular plant evolution, tracheids were the only type of conductive cells found in fossils of plant xylem tissues. [5] Ancestral tracheids did not contribute significantly to structural support, as can be seen in extant ferns. [6]