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Although its primary function is transport of sugars, phloem may also contain cells that have a mechanical support function. These are sclerenchyma cells which generally fall into two categories: fibres and sclereids. Both cell types have a secondary cell wall and are dead at maturity.
Cross section of celery stalk, showing vascular bundles, which include both phloem and xylem Detail of the vasculature of a bramble leaf Translocation in vascular plants. Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem ...
The cambium present between primary xylem and primary phloem is called the intrafascicular cambium (within vascular bundles). During secondary growth, cells of medullary rays, in a line (as seen in section; in three dimensions, it is a sheet) between neighbouring vascular bundles, become meristematic and form new interfascicular cambium ...
The phloem tissue arises from the cell facing outside and consists of sieve tubes and their companion cells. The function of phloem tissue is to distribute food from photosynthetic tissue to other tissues. The two tissues are separated by cambium, a tissue that divides to form xylem or phloem cells.
In vascular cambium, the primary phloem and xylem are produced by the apical meristem. After this initial development, secondary phloem and xylem are produced by the lateral meristem. The two are connected through a thin layer of parenchymal cells which are differentiated into the fascicular cambium.
Primarily, phloem carries dissolved food substances throughout the plant. This conduction system is composed of sieve-tube member and companion cells, that are without secondary walls. The parent cells of the vascular cambium produce both xylem and phloem. This usually also includes fibers, parenchyma and ray cells.
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Sieve elements are specialized cells that are important for the function of phloem, which is a highly organized tissue that transports organic compounds made during photosynthesis. Sieve elements are the major conducting cells in phloem. Conducting cells aid in transport of molecules especially for long-distance signaling.