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A fish's hypoxia tolerance can be represented in different ways. A commonly used representation is the critical O 2 tension (P crit), which is the lowest water O 2 tension (P O 2) at which a fish can maintain a stable O 2 consumption rate (M O 2). [2]
The concentration of oxygen in water is lower than air and it diffuses more slowly. In a litre of freshwater the oxygen content is 8 cm 3 per litre compared to 210 in the same volume of air. [7] Water is 777 times more dense than air and is 100 times more viscous. [7] Oxygen has a diffusion rate in air 10,000 times greater than in water. [7]
In some fish, capillary blood flows in the opposite direction to the water, causing countercurrent exchange. The muscles on the sides of the pharynx push the oxygen-depleted water out the gill openings. In bony fish, the pumping of oxygen-poor water is aided by a bone that surrounds the gills called the operculum. [6]
The oxygen in these tubes is renewed through the gills. In the larval dragonfly, the wall of the caudal end of the alimentary tract is richly supplied with tracheae as a rectal gill, and water pumped into and out of the rectum provides oxygen to the closed tracheae.
The gills push the oxygen-poor water out through openings in the sides of the pharynx. Fish from multiple groups can live out of the water for extended time periods. Amphibious fish such as the mudskipper can live and move about on land for up to several days, or live in stagnant or otherwise oxygen depleted water. Many such fish can breathe ...
The diet of the arapaima consists of fish, crustaceans, fruits, seeds, insects, and small land animals that walk near the shore. [18] The fish is an air breather, using its labyrinth organ, which is rich in blood vessels and opens into the fish's mouth, [19] an advantage in oxygen-deprived water that is often found in the Amazon River.
This organ allows labyrinth fish to take in oxygen directly from the air, instead of taking it from the water in which they reside through use of gills. The labyrinth organ helps the inhaled oxygen to be absorbed into the bloodstream. As a result, labyrinth fish can survive for a short period of time out of water, as they can inhale the air ...
Mudskippers are amphibious. When leaving the water and moving into a more dry environment on land, they are still able to breathe using water that is trapped inside their large gill chambers. They are also able to absorb oxygen from the lining of their mouth and throat, allowing them to stay out of water for long periods of time.