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In botany, a stoma (pl.: stomata, from Greek στόμα, "mouth"), also called a stomate (pl.: stomates), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange between the internal air spaces of the leaf and the atmosphere.
The stomata complex regulates the exchange of gases and water vapor between the outside air and the interior of the leaf. Typically, the stomata are more numerous over the abaxial (lower) epidermis of the leaf than the (adaxial) upper epidermis. An exception is floating leaves where most or all stomata are on the upper surface.
The pores or stomata of the epidermis open into substomatal chambers, which are connected to the intercellular air spaces between the spongy and palisade mesophyll cell, so that oxygen, carbon dioxide and water vapor can diffuse into and out of the leaf and access the mesophyll cells during respiration, photosynthesis and transpiration.
Guard cells are cells surrounding each stoma. They help to regulate the rate of transpiration by opening and closing the stomata. Light is the main trigger for the opening or closing. [citation needed] Each guard cell has a relatively thick and thinner cuticle [clarification needed] on the pore-side and a thin one opposite it. As water enters ...
The outside of the stem is covered with an epidermis, which is covered by a waterproof cuticle. The epidermis also may contain stomata for gas exchange and multicellular stem hairs called trichomes. A cortex consisting of hypodermis (collenchyma cells) and endodermis (starch containing cells) is present above the pericycle and vascular bundles.
They also separate stomata apart from each other as stomata have at least one pavement cell between each other. [4] They do not have a regular shape. Rather, their irregular shapes help them to interlock with each other like puzzle pieces to form a sturdy layer. [5]
A plant cuticle is a protecting film covering the outermost skin layer of leaves, young shoots and other aerial plant organs (aerial here meaning all plant parts not embedded in soil or other substrate) that have no periderm. The film consists of lipid and hydrocarbon polymers infused with wax, and is synthesized exclusively by the epidermal cells.
In citrus fruits, the flavedo constitutes the peripheral surface of the pericarp. It is composed of several cell layers that become progressively thicker in the internal part; the epidermic layer is covered with wax and contains few stomata, which in many cases are closed when the fruit is ripe.