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It forms a protective covering on the leaf vein and consists of one or more cell layers, usually parenchyma. Loosely-arranged mesophyll cells lie between the bundle sheath and the leaf surface. The Calvin cycle is confined to the chloroplasts of these bundle sheath cells in C 4 plants. C 2 plants also use a variation of this structure. [1]
D: Bundle sheath cell E: Stoma F: Vascular tissue 1. CO 2 is fixed to produce a four-carbon molecule (malate or aspartate). 2. The molecule exits the cell and enters the bundle sheath cells. 3. It is then broken down into CO 2 and pyruvate. CO 2 enters the Calvin cycle to produce carbohydrates. 4.
In the C4 pathway, a layer of mesophyll cells encircles bundle sheath cells that have large chloroplasts necessary for the Calvin cycle. A: Mesophyll Cell B: Chloroplast C: Vascular Tissue D: Bundle Sheath Cell E: Stroma F: Vascular Tissue: provides continuous source of water 1) Carbon is fixed to produce oxaloacetate by PEP carboxylase.
C 4 plants, in contrast, concentrate CO 2 spatially, with a RuBisCO reaction centre in a "bundle sheath cell" being inundated with CO 2. Due to the inactivity required by the CAM mechanism, C 4 carbon fixation has a greater efficiency in terms of PGA synthesis.
C 3 carbon fixation occurs in all plants as the first step of the Calvin–Benson cycle. (In C 4 and CAM plants, carbon dioxide is drawn out of malate and into this reaction rather than directly from the air.) Cross section of a C 3 plant, specifically of an Arabidopsis thaliana leaf. Vascular bundles shown.
2 concentrations in the Bundle Sheath are approximately 10–20 fold higher than the concentration in the mesophyll cells. [6] This ability to avoid photorespiration makes these plants more hardy than other plants in dry and hot environments, wherein stomata are closed and internal carbon dioxide levels are low.
The cells of the vascular cambium (F) divide to form phloem on the outside, located beneath the bundle cap (E), and xylem (D) on the inside. Most of the vascular cambium is here in vascular bundles (ovals of phloem and xylem together) but it is starting to join these up as at point F between the bundles.
As the plant grows, new vascular tissue differentiates in the growing tips of the plant. The new tissue is aligned with existing vascular tissue, maintaining its connection throughout the plant. The vascular tissue in plants is arranged in long, discrete strands called vascular bundles. These bundles include both xylem and phloem, as well as ...