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Conifers typically bear their microsporangia on microsporophylls aggregated into papery pollen strobili, and the ovules, are located on modified stem axes forming compound ovuliferous cone scales. Flowering plants contain microsporangia in the anthers of stamens (typically four microsporangia per anther) and megasporangia inside ovules inside ...
Encephalartos villosus microsporophylls with microsporangia. A microsporangium (pl. microsporangia) is a sporangium that produces microspores that give rise to male gametophytes when they germinate. Microsporangia occur in all vascular plants that have heterosporic life cycles, such as seed plants, spike mosses and the aquatic fern genus Azolla.
An anther and its filament together form a typical (or filantherous) stamen, part of the male floral organ. The typical anther is bilocular, i.e. it consists of two thecae. Each theca contains two microsporangia, also known as pollen sacs. The microsporangia produce the microspores, which for seed plants are known as pollen grains.
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The microsporangia of gymnosperms develop in pairs toward the bases of the scales, which are therefore called microsporophylls. Each of the microsporocytes in the microsporangia undergoes meiosis, producing four haploid microspores. These develop into pollen grains, each consisting of four cells and, in conifers, a pair of external air sacs.
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The Doomsday Clock is seen at 89 seconds to midnight, the closest the clock has ever been to midnight in its 78-year history. / Credit: SAUL LOEB/AFP via Getty Images
In heterosporous plants, sporophylls (whether they are microphylls or megaphylls) bear either megasporangia and thus are called megasporophylls, or microsporangia and are called microsporophylls. The overlap of the prefixes and roots makes these terms a particularly confusing subset of botanical nomenclature.