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Scutes on an alligator foot A scute ( / s k j uː t / ⓘ ) or scutum ( Latin : scutum ; plural: scuta " shield ") is a bony external plate or scale overlaid with horn, as on the shell of a turtle , the skin of crocodilians , and the feet of birds .
In the center of the carapace are five vertebral scutes and out from these are four pairs of costal scutes. Around the edge of the shell are 12 pairs of marginal scutes. All these scutes are aligned so that for the most part the sutures between the bones are in the middle of the scutes above.
Typically, 11 or 12 pairs of marginal scutes rim the carapace. [8] Five vertebral scutes run down the carapace's midline, while five pairs of costal scutes border them. [21] The nuchal scute is located at the base of the head. [21] The carapace connects to the plastron by three pairs of inframarginal scutes forming the bridge of the shell. [21]
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Snake scales may be modified to form fringes, as in the case of the eyelash bush viper, Atheris ceratophora, or rattles as in the case of the rattlesnakes of North America. [ 8 ] Certain primitive snakes such as boas , pythons and certain advanced snakes such as vipers have small scales arranged irregularly on the head.
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The articulations with neighboring scutes are located on the ventral side, and the dorsal surface is strongly sculptured while the ventral surface of wings is smooth and contains many small openings. "The scutes have very wide wings and cover practically the whole upper part of the trunk. The tail scutes are narrower, the last of which lack wings.
Squamata and Rhynchocephalia form the subclass Lepidosauria, which is the sister group to the Archosauria, the clade that contains crocodiles and birds, and their extinct relatives. Fossils of rhynchocephalians first appear in the Early Triassic , meaning that the lineage leading to squamates must have also existed at the time.