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The butterflies form the clade Rhopalocera, which is composed of three superfamilies: Hedyloidea (the moth butterfly family Hedylidae), the Hesperioidea (the skipper family Hesperiidae), and the Papilionoidea (the true butterfly families Papilionidae, Pieridae, Nymphalidae, Lycaenidae, and Riodinidae). All of these families are monophyletic.
They evolved about the time that flowering plants underwent an expansive adaptive radiation in the mid-Cretaceous, and the Gelechioidea that evolved at this time also have great diversity. Whether the processes involved co-evolution or sequential evolution, the diversity of the Lepidoptera and the angiosperms increased together.
The evolution of birds began in the Jurassic Period, with the earliest birds derived from a clade of theropod dinosaurs named Paraves. [1] Birds are categorized as a biological class, Aves. For more than a century, the small theropod dinosaur Archaeopteryx lithographica from the Late Jurassic period was considered to have been the earliest bird.
Butterflies evolved from moths, so while the butterflies are monophyletic (forming a single clade), the moths are not. The oldest known butterfly is Protocoeliades kristenseni from the Palaeocene aged Fur Formation of Denmark, approximately 55 million years old, which belongs to the family Hesperiidae (skippers). [ 9 ]
Owl butterflies are very large, 65–200 mm (2.6–7.9 in), and fly only a few meters at a time, so avian predators have little difficulty in following them to their settling place. However, the butterflies preferentially fly in dusk, when few avian predators are around. [ 1 ]
A turning point came in the early twentieth century with the writings of Gerhard Heilmann of Denmark.An artist by trade, Heilmann had a scholarly interest in birds and from 1913 to 1916, expanding on earlier work by Othenio Abel, [12] published the results of his research in several parts, dealing with the anatomy, embryology, behavior, paleontology, and evolution of birds. [13]
Lepidoptera (/ ˌ l ɛ p ɪ ˈ d ɒ p t ər ə / LEP-ih-DOP-tər-ə) or lepidopterans is an order of winged insects which includes butterflies and moths.About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organisms, [1] [2] making it the second largest insect order (behind Coleoptera) with 126 families [3] and 46 superfamilies ...
A Lepidoptera specimen drawer in a museum collection in Poland Another Lepidoptera specimen drawer in a museum collection in Poland. Lepidopterology (from Ancient Greek λεπίδος (lepídos) ' scale ' πτερόν (pterón) ' wing ' and -λογία [1]) is a branch of entomology concerning the scientific study of moths and the two superfamilies of butterflies.