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Pollen tube elongation is an integral stage in the plant life cycle. The pollen tube acts as a conduit to transport the male gamete cells from the pollen grain—either from the stigma (in flowering plants) to the ovules at the base of the pistil or directly through ovule tissue in some gymnosperms.
Sieve cells are long, conducting cells in the phloem that do not form sieve tubes. The major difference between sieve cells and sieve tube members is the lack of sieve plates in sieve cells. [1] They have a very narrow diameter and tend to be longer in length than sieve tube elements as they are generally associated with albuminous cells. [4]
Microtubule and tubulin metrics [1]. Microtubules are polymers of tubulin that form part of the cytoskeleton and provide structure and shape to eukaryotic cells. Microtubules can be as long as 50 micrometres, as wide as 23 to 27 nm [2] and have an inner diameter between 11 and 15 nm. [3]
The microtubule-organizing center (MTOC) is a structure found in eukaryotic cells from which microtubules emerge. MTOCs have two main functions: the organization of eukaryotic flagella and cilia and the organization of the mitotic and meiotic spindle apparatus, which separate the chromosomes during cell division.
Once the pollen lands on the stigma of a receptive flower, it germinates and starts growing a pollen tube through the carpel. Before the tube reaches the ovule, the nucleus of the generative cell in the pollen grain divides and gives rise to two sperm nuclei, which are then discharged through the tube into the ovule for fertilization. [34]
An endosperm is formed after the two sperm nuclei inside a pollen grain reach the interior of a female gametophyte or megagametophyte, also called the embryonic sac.One sperm nucleus fertilizes the egg cell, forming a zygote, while the other sperm nucleus usually fuses with the binucleate central cell, forming a primary endosperm cell (its nucleus is often called the triple fusion nucleus).
Equally as important in the function of endocytosis is the structure known as the cytopharynx. The cytopharynx is a long, tube-like structure that forms the invagination associated with the cytostome. While the shape of the cytopharynx is not constant, it is typically directed towards the posterior of the cell, often hooking around a central ...
All of the cellular functions of a sieve-tube element are carried out by the (much smaller) companion cell, a typical nucleate plant cell except the companion cell usually has a larger number of ribosomes and mitochondria. The dense cytoplasm of a companion cell is connected to the sieve-tube element by plasmodesmata. [10]