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The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
A speculatively rooted tree for RNA genes, showing major branches Bacteria, Archaea, and Eukaryota The three-domain tree and the eocyte hypothesis (two-domain tree), 2008. [7] Phylogenetic tree showing the relationship between the eukaryotes and other forms of life, 2006. [8] Eukaryotes are colored red, archaea green, and bacteria blue.
The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. [1] [2] [3] It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya. [4]
Woese argued that the bacteria, archaea, and eukaryotes represent separate lines of descent that diverged early on from an ancestral colony of organisms. [ 45 ] [ 46 ] However, a few biologists argue that the Archaea and Eukaryota arose from a group of bacteria. [ 47 ]
In some phylogenetic trees based upon different gene/protein sequences of prokaryotic homologs, the archaeal homologs are more closely related to those of gram-positive bacteria. [82] Archaea and gram-positive bacteria also share conserved indels in a number of important proteins, such as Hsp70 and glutamine synthetase I; [82] [84] but the ...
In 1977, Carl Woese and colleagues proposed the fundamental subdivision of the prokaryotes into the Eubacteria (later called the Bacteria) and Archaebacteria (later called the Archaea), based on ribosomal RNA structure; [15] this would later lead to the proposal of three "domains" of life, of Bacteria, Archaea, and Eukaryota. [5]
A major step forward in the study of bacteria came in 1977 when Carl Woese recognised that archaea have a separate line of evolutionary descent from bacteria. [263] This new phylogenetic taxonomy depended on the sequencing of 16S ribosomal RNA and divided prokaryotes into two evolutionary domains, as part of the three-domain system .
Archaea share this defining feature with the bacteria with which they were once grouped. In 1990 the microbiologist Woese proposed the three-domain system that divided living things into bacteria, archaea and eukaryotes, [42] and thereby split the prokaryote domain. Archaea differ from bacteria in both their genetics and biochemistry.