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The Boss 429 was produced in limited numbers in 1969 and 1970. Its origin is twofold: allow Ford to homologate its new Ford 385 engine -based semi-hemispherical 429 V8 for NASCAR racing, and to provide a big block alternative to the 5.0 L and under Trans Am Racing bred small block Boss 302 Mustang .
At mid-year, Ford offered a slightly detuned Boss 351 engine, which could be ordered with any model. A total of 398 Mustangs were built with the drag race-oriented R code engine and were designated as the 351 HO. Mandatory options were the top loader 4-speed, competition N case rear end (427, 428, 429, Boss 351, 351HO), and air conditioning delete.
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Later in the season, the Boss 429 engine was used by many of the teams, after it had finally been declared "officially homologated" by NASCAR president Bill France. The Boss 429 engine was homologated in the 1969 Ford Mustang Boss 429. In a very unusual move, Ford homologated the engine separately from the car in which it was to race.
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A little-known and extremely rare offering in 1969 was the Rio Grande. Available on special order, this was essentially a GT in the so-called "Grabber" colors of "Wimbledon White", "Poppy Red" or "Calypso Coral", partially blacked-out hood with scoop, side stripes, bed rails, vinyl top, and unique "Ford Ranchero Rio Grande" wheel centers.
For 1970, the two-barrel 351 remained the base engine, with the four-barrel engine replaced by a 351 cubic-inch "Cleveland" V8, rated at 300 hp (224 kW; 304 PS). While unchanged in displacement, this engine was an all-new design. The 390 was dropped, with the 428 Cobra Jet remaining alongside the Boss 302 as the high-performance engine options.