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Single heterocysts develop about every 9-15 cells, producing a one-dimensional pattern along the filament. The interval between heterocysts remains approximately constant even though the cells in the filament are dividing. The bacterial filament can be seen as a multicellular organism with two distinct yet interdependent cell types.
The cells are bacteria and thus have no nucleus nor internal membrane system. To multiply, they form two new cells when they divide by binary fission . Along the trichomes, larger specialist nitrogen-fixing cells called heterocysts occur between the ordinary cells.
Intercalary located akinete of Dolichospermum smithii Terminally located akinete of Gloeotrichia. An akinete is an enveloped, thick-walled, non-motile, dormant cell formed by filamentous, heterocyst-forming cyanobacteria under the order Nostocales and Stigonematales.
Their cell size varies from less than 1 μm in diameter (picocyanobacteria) up to 100 μm (some tropical forms in the genus Oscillatoria) [39] [40] [41] Filamentous forms exhibit functional cell differentiation such as heterocysts (for nitrogen fixation), akinetes (resting stage cells), and hormogonia (reproductive, motile filaments). These ...
Nostoc, also known as star jelly, troll's butter, spit of moon, fallen star, witch's butter (not to be confused with the fungi commonly known as witches' butter), and witch's jelly, is the most common genus of cyanobacteria found in a variety of both aquatic and terrestrial environments that may form colonies composed of filaments of moniliform cells in a gelatinous sheath of polysaccharides. [1]
Nitrogenase, sequestered within these cells, transforms dinitrogen into ammonia at the expense of ATP and reductant—both generated by carbohydrate metabolism, a process supplemented, in the light, by the activity of PS I. Carbohydrate, probably in the form of glucose, is synthesized in vegetative cells and moves into heterocysts.
The appearance of cell differentiation was a great evolutionary leap; marking cyanobacteria as one of the first multicellular organisms on Earth. [2] On the A. circinalis filament, the most numerous structures are vegetative cells, responsible for the photosynthesis of high-energy sugars from environmental carbon, water, and sunlight. The ...
The classic taxonomic criterion has been the cell morphology and the plane of cell division. In Pleurocapsales, the cells have the ability to form internal spores (baeocytes). The rest of the sections include filamentous species. In Oscillatoriales, the cells are uniseriately arranged and do not form specialized cells (akinetes and heterocysts ...