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Agnatha (/ ˈ æ ɡ n ə θ ə, æ ɡ ˈ n eɪ θ ə /; [3] from Ancient Greek ἀ-(a-) 'without' and γνάθος (gnáthos) 'jaws') is a paraphyletic infraphylum [4] of non-gnathostome vertebrates, or jawless fish, in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and extinct (conodonts, anaspids, and ostracoderms, among others).
Cyclostomi, often referred to as Cyclostomata / s ɪ k l oʊ ˈ s t ɒ m ə t ə /, is a group of vertebrates that comprises the living jawless fishes: the lampreys and hagfishes.Both groups have jawless mouths with horny epidermal structures that function as teeth called ceratodontes, and branchial arches that are internally positioned instead of external as in the related jawed fishes. [1]
Recent molecular and morphological phylogenetic studies place lampreys and hagfish in the infraphylum Agnatha or Agnathostomata (both meaning without jaws). The other vertebrate infraphylum is Gnathostomata (jawed mouths) and includes the classes Chondrichthyes (sharks), Osteichthyes (bony fishes), Amphibia, Reptilia, Aves, and Mammalia.
Infraphylum Agnatha or Cephalaspidomorphi (lampreys and other jawless fishes) Superclass † Anaspidomorphi (anaspids and relatives) Infraphylum Gnathostomata (vertebrates with jaws)
Gnathostomata is traditionally a infraphylum, broken into three top-level groupings: Chondrichthyes, or the cartilaginous fish; Placodermi, an extinct grade of armored fish; and Teleostomi, which includes the familiar classes of bony fish, birds, mammals, reptiles, and amphibians. Some classification systems have used the term Amphirhina.
Jawless fish belong to the superclass Agnatha in the phylum Chordata, subphylum Vertebrata. Agnatha means 'un-jawed, without jaws' (from Ancient Greek). [11] It excludes all vertebrates with jaws, known as gnathostomes. Although a minor element of modern marine fauna, jawless fish were prominent among the early fish in the early Paleozoic.
Other elements of the skull, however, may be reduced; there is little cheek region behind the enlarged orbits, and little, if any bone in between them. The upper jaw is often formed largely from the premaxilla , with the maxilla itself located further back, and an additional bone, the symplectic, linking the jaw to the rest of the cranium.
Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish ().According to FishBase, 33,400 species of fish had been described as of October 2016, with approximately 250 new species described each year.