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Pollen tubes are an excellent model for the understanding of plant cell behavior. [19] They are easily cultivated in vitro and have a very dynamic cytoskeleton that polymerizes at very high rates, providing the pollen tube with interesting mechanical properties. [20] The pollen tube has an unusual kind of growth; it extends exclusively at its apex.
If the pollen is compatible it will germinate and begin to grow. [5] The ovary releases chemicals that stimulates a positive chemotropic response from the developing pollen tube. [6] In response the tube develops a defined tip growth area that promotes directional growth and elongation of the pollen tube due to a calcium gradient. [5]
The anthers are joined forming a tube around the style in which the pollen is released, and the style then grows through this tube, pushing out or taking up the pollen (with variably developed hairs) and presenting it to floral visitors, after which the stigmas become receptive (i.e., with a plunger or brush pollination mechanism).
The pollen is carried to the pistil of another flower, by wind or animal pollinators, and deposited on the stigma. As the pollen grain germinates, the tube cell produces the pollen tube, which elongates and extends down the long style of the carpel and into the ovary, where its sperm cells are released in the megagametophyte.
Floral diagram of Anagallis arvensis. [1]: 307 The dot represents the main axis, green structure below is the subtending bract.Calyx (green arcs) consists of five free sepals; corolla (red arcs) consists of five fused petals.
The style is a hollow tube in some plants, such as lilies, or has transmitting tissue through which the pollen tubes grow. [15] The stigma (from Ancient Greek στίγμα, stigma, meaning mark or puncture) is usually found at the tip of the style, the portion of the carpel(s) that receives pollen (male gametophytes). It is commonly sticky or ...
The pollen grains, which are the male gametophytes, are reduced to only a few cells (just three cells in many cases). Here the notion of two generations is less obvious; as Bateman & Dimichele say "sporophyte and gametophyte effectively function as a single organism". [8] The alternative term 'alternation of phases' may then be more appropriate ...
Gnetophyta may have 2 or 3 celled pollen grains depending on the species, and Coniferophyta pollen grains vary greatly ranging from single celled to 40 celled. [11] [10] One of these cells is typically a germ cell and other cells may consist of a single tube cell which grows to form the pollen tube, sterile cells, and/or prothallial cells which ...