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Stoma in a tomato leaf shown via colorized scanning electron microscope image A stoma in horizontal cross section The underside of a leaf. In this species (Tradescantia zebrina) the guard cells of the stomata are green because they contain chlorophyll while the epidermal cells are chlorophyll-free and contain red pigments.
Photosynthesis depends on the diffusion of carbon dioxide (CO 2) from the air through the stomata into the mesophyll tissues. Oxygen (O 2), produced as a byproduct of photosynthesis, exits the plant via the stomata. When the stomata are open, water is lost by evaporation and must be replaced via the transpiration stream, with water taken up by ...
The pores or stomata of the epidermis open into substomatal chambers, which are connected to the intercellular air spaces between the spongy and palisade mesophyll cell, so that oxygen, carbon dioxide and water vapor can diffuse into and out of the leaf and access the mesophyll cells during respiration, photosynthesis and transpiration.
The stomata complex regulates the exchange of gases and water vapor between the outside air and the interior of the leaf. Typically, the stomata are more numerous over the abaxial (lower) epidermis of the leaf than the (adaxial) upper epidermis. An exception is floating leaves where most or all stomata are on the upper surface.
[4] [6] During the following day, stomata are closed, malate is decarboxylated, and CO 2 is fixed by RuBisCO. This process alone is similar to that of C4 plants and yields characteristic C4 fractionation values of approximately -11‰. [6] However, in the afternoon, CAM plants may open their stomata and perform C3 photosynthesis. [6]
The pineapple is an example of a CAM plant.. Crassulacean acid metabolism, also known as CAM photosynthesis, is a carbon fixation pathway that evolved in some plants as an adaptation to arid conditions [1] that allows a plant to photosynthesize during the day, but only exchange gases at night.
It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, engages in photosynthesis, stores nutrients, and produces new living tissue. [1] The stem can also be called the culm, halm, haulm, stalk, or thyrsus. The stem is normally divided into nodes and internodes: [2]
Many cacti conduct photosynthesis in succulent stems, rather than leaves, so the surface area of the shoot is very low. Many desert plants have a special type of photosynthesis, termed crassulacean acid metabolism or CAM photosynthesis, in which the stomata are closed during the day and open at night when transpiration will be lower. [14]