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The order Ichthyostegalia was erected for Ichthyostega, and contained until the 1980s only three genera (Ichthyostega, Acanthostega and Tulerpeton).While Ichthyostegalia in principle contain the most basal of animals with toes rather than fins, Clack and Ahlberg uses it for all finds more advanced than Tiktaalik (the closest relative of tetrapods known to have retained paired fins rather than ...
Ichthyostega (from Greek: ἰχθῦς ikhthûs, 'fish' and Greek: στέγη stégē, 'roof') is an extinct genus of limbed tetrapodomorphs from the Late Devonian of what is now Greenland. It was among the earliest four-limbed vertebrates ever in the fossil record and was one of the first with weight-bearing adaptations for terrestrial locomotion.
Tiktaalik also lacked a characteristic most fishes have—bony plates in the gill area that restrict lateral head movement. This makes Tiktaalik the earliest-known fish to have a neck, with the pectoral (shoulder) girdle separate from the skull. This would give the creature more freedom in hunting prey on land or in the shallows.
In the tetrapod and higher clades from the lower-middle Famennian there are several defining changes on the basis of anatomy of Ichthyostega, Tulerpeton, and Acanthostega. In the cranium, there is a stapes derived from the hyomandibular of fishes; a single bilateral pair of nasal bones, and a fenestra ovalis in the otic capsule of the braincase ...
In Late Devonian vertebrate speciation, descendants of pelagic lobe-finned fish such as Eusthenopteron exhibited a sequence of adaptations: Panderichthys, suited to muddy shallows; Tiktaalik with limb-like fins that could take it onto land; early tetrapods in weed-filled swamps, such as Acanthostega, which had feet with eight digits, and Ichthyostega, which had limbs.
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Tiktaalik, a tetrapodomorph with wrists, straddles the fish-tetrapod divide. The Stem Tetrapoda are a cladistically defined group, consisting of all animals more closely related to extant four-legged vertebrates than to their closest extant relatives (the lungfish), but excluding the crown group Tetrapoda.