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Subfamily Pinoideae : cones are biennial, rarely triennial, with each year's scale-growth distinct, forming an umbo on each scale, the cone scale base is broad, concealing the seeds fully from abaxial (below the phloem vessels) view, the seed is without resin vesicles, the seed wing holds the seed in a pair of claws, leaves have primary ...
The cones thus grow over a two-year (26-month) cycle, so that newer green and older, seed-bearing or open brown cones are on the tree at the same time. Open cone with empty pine nuts. The seed cones open to 6–9 cm (2 + 1 ⁄ 4 – 3 + 1 ⁄ 2 in) broad when mature, holding the seeds on the scales after opening.
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A mature female big-cone pine (Pinus coulteri) cone, the heaviest pine cone A young female cone on a Norway spruce (Picea abies) Immature male cones of Swiss pine (Pinus cembra) A conifer cone, or in formal botanical usage a strobilus, pl.: strobili, is a seed-bearing organ on gymnosperm plants, especially in conifers and cycads.