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Mona tells the other girls that she met Leslie in Radley. This enforces the girls theory that Leslie is "A". At the end of the episode Mona is seen on the phone to Leslie saying that she failed to keep her time in Radley a secret. Leslie said that Mona is useless and messes everything up like usual. Mona appears at the ending of "No Stone ...
Mona is out of Radley and back at Rosewood High. The Liars do not trust Mona. Someone plants a cow brain in Mona's locker kept up with a knife. Mona is appalled and whispers something to Lucas. The Liars worry that people will think they were involved, including Mona, who instantly assures them she believes them, and wants to earn back their trust.
Mona locates Alison's phone at the Carissimi Group building, and the girls head to the company's headquarters. There, Spencer breaks into a restricted room after Sara remembers a specific date meaningful to Charles, which is the code for unlocking the door. While the girls enter the room, Sara refuses to accompany them.
After Mona is revealed to be "A", Wren develops a friendship with Hanna, as he volunteers at Radley Sanitarium where Hanna goes to visit Mona there on a regular basis. Eventually, the board at Radley threatens to transfer Mona to New York, and Wren helps Hanna testify to the board and gets her to remain at Radley.
Though Mona survives, she is sent to Radley Sanitarium for medical assistance. While in the psychiatric hospital, Mona takes up a partnership with a new accomplice that starts off the second game. After this, Mona became another henchman in the "A" hierarchy, obeying the orders of Big A, known as Red Coat, the captain of the A-Team.
Mona looks through the Radley files and discovers that Alison knew about the affair and lured Bethany to Rosewood. Mona calls Aria and tells her the information, and that Alison must be 'A'. However, before the girls arrive to Mona's house, "A", with blonde hair cascading out from their hoodie, attacks Mona.
The next day, the Liars and Mona are at the Radley, talking about what happened the night before. Surprising everyone, Jenna Marshall ( Tammin Sursok ) suddenly appears; she is surprised when she learns that the girls are there, and reveals that she is back to Rosewood to celebrate Toby and Yvonne's engagement.
Meanwhile, Mona later returns to Rosewood, heavily questioned by the police about her actions while her friendship with Lesli is strained because of her involvement with Mona's actions. It is later revealed that Lesli was a former patient at Radley, and was roommates with Bethany Young, leading the girls to suspect that Lesli may be 'A'.