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[3]: 354 [4] [5] Differences in behavior or biology that inhibit formation of hybrid zygotes are termed prezygotic isolation. Reinforcement can be shown to be occurring (or to have occurred in the past) by measuring the strength of prezygotic isolation in a sympatric population in comparison to an allopatric population of the same species.
The types of barriers that can cause this isolation include: different habitats, physical barriers, and a difference in the time of sexual maturity or flowering. [6] [7] An example of the ecological or habitat differences that impede the meeting of potential pairs occurs in two fish species of the family Gasterosteidae (sticklebacks).
For example: If population has the prezygotic isolating allele and the high fitness, post-zygotic alleles and ; and population has the prezygotic allele a and the high fitness, post-zygotic alleles and , both and genotypes will experience recombination in the face of gene flow. Somehow, the populations must be maintained.
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A major aspect of speciation research is to determine the nature of the barriers that inhibit reproduction. Botanists often consider the zoological classifications of prezygotic and postzygotic barriers as inadequate. [259] The examples provided below give insight into the process of speciation.