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The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. [1] [2] [3] It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya. [4]
A speculatively rooted tree for RNA genes, showing major branches Bacteria, Archaea, and Eukaryota The three-domain tree and the eocyte hypothesis (two-domain tree), 2008. [7] Phylogenetic tree showing the relationship between the eukaryotes and other forms of life, 2006. [8] Eukaryotes are colored red, archaea green, and bacteria blue.
Archaea (/ ɑːr ˈ k iː ə / ⓘ ar-KEE-ə) is a domain of organisms.Traditionally, Archaea only included its prokaryotic members, but since this has been found to be paraphyletic, as eukaryotes are now known to have evolved from archaea.
A major step forward in the study of bacteria came in 1977 when Carl Woese recognised that archaea have a separate line of evolutionary descent from bacteria. [263] This new phylogenetic taxonomy depended on the sequencing of 16S ribosomal RNA and divided prokaryotes into two evolutionary domains, as part of the three-domain system .
Woese argued that the bacteria, archaea, and eukaryotes represent separate lines of descent that diverged early on from an ancestral colony of organisms. [ 45 ] [ 46 ] However, a few biologists argue that the Archaea and Eukaryota arose from a group of bacteria. [ 47 ]
The combined domains of archaea and bacteria make up the most diverse and abundant group of organisms on Earth and inhabit practically all environments where the temperature is below +140 °C (284 °F). They are found in water, soil, air, as the microbiome of an organism, hot springs and even deep beneath the Earth's crust in rocks. [48]
Archaea and bacteria are structurally similar even though they are not closely related in the tree of life. The shapes of both bacteria and archaea cells vary from a spherical shape known as coccus or a rod-shape known as bacillus. They are also related with no internal membrane and a cell wall that assists the cell maintaining its shape.