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The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
Archaea were first classified separately from bacteria in 1977 by Carl Woese and George E. Fox, based on their ribosomal RNA (rRNA) genes. [14] (At that time only the methanogens were known). They called these groups the Urkingdoms of Archaebacteria and Eubacteria, though other researchers treated them as kingdoms or subkingdoms.
The tree of life. Two domains of life are Bacteria (top branches) and Archaea (bottom branches, including eukaryotes). The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea.
Phylogenetic tree showing the relationship between the archaea and other forms of life. Eukaryotes are colored red, archaea green and bacteria blue. Adapted from Ciccarelli et al. [44] Woese argued that the bacteria, archaea, and eukaryotes represent separate lines of descent that diverged early on from an ancestral colony of organisms.
A speculatively rooted tree for RNA genes, showing major branches Bacteria, Archaea, and Eukaryota The three-domain tree and the eocyte hypothesis (two-domain tree), 2008. [7] Phylogenetic tree showing the relationship between the eukaryotes and other forms of life, 2006. [8] Eukaryotes are colored red, archaea green, and bacteria blue.
However, molecular systematics showed prokaryotic life to consist of two separate domains, originally called Eubacteria and Archaebacteria, but now called Bacteria and Archaea that evolved independently from an ancient common ancestor. [5] The archaea and eukaryotes are more closely related to each other than either is to the bacteria.
Metagenomic analyses recover a two-domain system with the domains Archaea and Bacteria; in this view of the tree of life, Eukaryotes are derived from Archaea. [ 57 ] [ 58 ] [ 59 ] With the later gene pool of LUCA's descendants, sharing a common framework of the AT/GC rule and the standard twenty amino acids, horizontal gene transfer would have ...
In 1977, Carl Woese and colleagues proposed the fundamental subdivision of the prokaryotes into the Eubacteria (later called the Bacteria) and Archaebacteria (later called the Archaea), based on ribosomal RNA structure; [15] this would later lead to the proposal of three "domains" of life, of Bacteria, Archaea, and Eukaryota. [5]