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In land plants, the plastids that contain chlorophyll can perform photosynthesis, thereby creating internal chemical energy from external sunlight energy while capturing carbon from Earth's atmosphere and furnishing the atmosphere with life-giving oxygen. These are the chlorophyll-plastids—and they are named chloroplasts; (see top graphic).
[3] [4] [5] This group contains the carbohydrate paramylon. Euglenids split from other Euglenozoa (a larger group of flagellates) more than a billion years ago. The plastids (membranous organelles) in all extant photosynthetic species result from secondary endosymbiosis between a euglenid and a green alga.
The alternative to monophyly is serial endosymbiosis, meaning that the "chromists" acquired their plastids from each other instead of inheriting them from a single common ancestor. Thus the phylogeny of the distinctive plastids, which are agreed to have a common origin in the rhodophytes, is different from the phylogeny of the host cells. [3]
A diagram showing the different types of plastid. Amyloplasts are thought to play a vital role in gravitropism.Statoliths, a specialized starch-accumulating amyloplast, are denser than cytoplasm, and are able to settle to the bottom of the gravity-sensing cell, called a statocyte. [5]
Photosynthetic organisms with plastids of different origin (such as brown algae) do not belong to the Archaeplastida. The archaeplastidans fall into two main evolutionary lines. The red algae are pigmented with chlorophyll a and phycobiliproteins, like most cyanobacteria, and accumulate starch outside the chloroplasts.
Kinetoplastida (or Kinetoplastea, as a class) is a group of flagellated protists belonging to the phylum Euglenozoa, [3] [4] and characterised by the presence of a distinctive organelle called the kinetoplast (hence the name), a granule containing a large mass of DNA.
Representative genera (examples) Description Discoba or JEH or Eozoa: Tsukubea: Tsukubamonas: Euglenozoa: Euglena, Trypanosoma: Many important parasites, one large group with plastids (chloroplasts) Heterolobosea (Percolozoa) Naegleria, Acrasis: Most alternate between flagellate and amoeboid forms Jakobea: Jakoba, Reclinomonas
The Apicomplexa (also called Apicomplexia; single: apicomplexan) are organisms of a large phylum of mainly parasitic alveolates.Most possess a unique form of organelle structure that comprises a type of non-photosynthetic plastid called an apicoplast—with an apical complex membrane.