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The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
This led to the conclusion that Archaea and Eukarya shared a common ancestor more recent than Eukarya and Bacteria. [73] The development of the nucleus occurred after the split between Bacteria and this common ancestor. [73] [2] One property unique to archaea is the abundant use of ether-linked lipids in their cell membranes.
Metagenomic analyses recover a two-domain system with the domains Archaea and Bacteria; in this view of the tree of life, Eukaryotes are derived from Archaea. [ 57 ] [ 58 ] [ 59 ] With the later gene pool of LUCA's descendants, sharing a common framework of the AT/GC rule and the standard twenty amino acids, horizontal gene transfer would have ...
The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. [1] [2] [3] It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya. [4]
The eukaryotes seemingly emerged within the Asgard archaea, and are closely related to the Heimdallarchaeia. [5] This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as flagellated cells.
Cellular life forms can be divided into prokaryotes and eukaryotes. Prokaryotes are bacteria or archaea, and the diagram shows some (clickable) parts shared by both. But bacteria and archaea also have fundamental differences, as indicated by their placement in different domains.
Archaea share this defining feature with the bacteria with which they were once grouped. In 1990 the microbiologist Woese proposed the three-domain system that divided living things into bacteria, archaea and eukaryotes, [42] and thereby split the prokaryote domain. Archaea differ from bacteria in both their genetics and biochemistry.
However, molecular systematics showed prokaryotic life to consist of two separate domains, originally called Eubacteria and Archaebacteria, but now called Bacteria and Archaea that evolved independently from an ancient common ancestor. [5] The archaea and eukaryotes are more closely related to each other than either is to the bacteria.