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By contrast, the envelope of gram-negative bacteria is more complex and consists (from inside to outside) of the cytoplasmic membrane, a thin layer of peptidoglycan, and an additional outer membrane, also called the lipopolysaccharide layer. Other bacterial cell surface structures range from disorganised slime layers to highly structured capsules.
Navicula diatoms are highly motile and move through a gliding movement [3] [4] [5] This is done through excretion of extracellular polymeric substances (EPS). One form of EPS surrounds the outside of the cell and another is excreted through a slit in the frustule called a raphe, allowing the cell to glide along a track.
Cells are rectangular in girdle view (when in colonies), and lanceolate in valve view. Raphe system is slightly keeled and runs from pole to pole. Two large plate-like chloroplasts are present, one near each end of the cell. The nucleus is located centrally. Cells are yellow-brown in colour.
The circular muscle layer prevents food from travelling backward and the longitudinal layer shortens the tract. The thickness of the muscular layer varies in each part of the tract: In the colon, for example, the muscular layer is much thicker because the faeces are large and heavy and require more force to push along.
Bacillaria paxillifer was originally described under the name Vibrio paxillifer by Otto Frederick Müller in 1786. It is the first diatom species known to be described. [5] It was separately described two years later (1788) by Johann Friedrich Gmelin as Bacillaria paradoxa.
A frustule is the hard and porous cell wall or external layer of diatoms. The frustule is composed almost purely of silica, made from silicic acid, and is coated with a layer of organic substance, which was referred to in the early literature on diatoms as pectin, a fiber most commonly found in cell walls of plants.
For many years the diatoms—treated either as a class (Bacillariophyceae) or a phylum (Bacillariophyta)—were divided into just 2 orders, corresponding to the centric and the pennate diatoms (Centrales and Pennales; alternative names Biddulphiales and Bacillariales, as used e.g. in Lee, 1989). [9]
Examples range from the propulsion of single cells such as the swimming of spermatozoa to the transport of fluid along a stationary layer of cells such as in a respiratory tract. Though eukaryotic flagella and motile cilia are ultrastructurally identical, the beating pattern of the two organelles can be different.