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The iris dilator muscle (pupil dilator muscle, pupillary dilator, radial muscle of iris, radiating fibers), is a smooth muscle [2] of the eye, running radially in the iris and therefore fit as a dilator. The pupillary dilator consists of a spokelike arrangement of modified contractile cells called myoepithelial cells.
The pupillary dilator muscle increases the pupillary diameter and it is arranged radially, but the pupillary sphincter muscle is responsible for the constriction of the pupil's diameter and it encircles the pupil. [2] The pupillary dilation is also called mydriasis, and the constriction of the pupil is also called miosis. [1]
The pupil constricts when the iris sphincter muscle is stimulated and contracts. In humans, it functions to constrict the pupil in bright light (pupillary light reflex) or during accommodation. [citation needed] In lower animals, the muscle cells themselves are photosensitive causing iris action without brain input. [5]
The iris consists of two layers: the front pigmented fibrovascular layer known as a stroma and, behind the stroma, pigmented epithelial cells.. The stroma is connected to a sphincter muscle (sphincter pupillae), which contracts the pupil in a circular motion, and a set of dilator muscles (dilator pupillae), which pull the iris radially to enlarge the pupil, pulling it in folds.
An area termed the limbus connects the cornea and sclera. The iris is the pigmented circular structure concentrically surrounding the centre of the eye, the pupil, which appears to be black. The size of the pupil, which controls the amount of light entering the eye, is adjusted by the iris' dilator and sphincter muscles.
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There are two types of muscle that control the size of the iris: the iris sphincter, composed of circularly arranged muscle fibers, and the iris dilator, composed of radially arranged muscle fibers. The sphincter is controlled by nerves of the parasympathetic nervous system, and the dilator by the sympathetic nervous system.
Structure of the iris and surrounding parts showing the stroma of iris (stroma iridis). The stroma connects to a sphincter muscle ( sphincter pupillae ), which contracts the pupil in a circular motion, and a set of dilator muscles ( dilator pupillae ) which pull the iris radially to enlarge the pupil, pulling it in folds.