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Lillie (Japanese: リーリエ, Hepburn: Rīrie) is a character in Nintendo and Game Freak's Pokémon franchise, introduced in the 2016 video games Pokémon Sun and Moon, being a central character in the story, and has appeared in numerous Pokémon media.
Lusamine (ルザミーネ) is the president of the Aether Foundation, the mother of Lillie and Gladion, and the true villain of the original Sun and Moon games. She is obsessed with the Ultra Beasts, specifically Nihilego , having fallen under their influence; Gladion speculates her fixation started after her husband Mohn vanished while ...
Ash, Gladion, Lillie, and Lusamine go after Faba, arriving just as a panicked Nebby transforms into a chrysalis and the Ultra Wormhole begins to open! The Ultra Beast known as UB-01 Symbiont emerges and attacks Lillie — but Lusamine throws herself in front of her daughter, and the Ultra Beast engulfs her instead and takes her back through the ...
Faba is a member of the Aether Foundation, a charitable organization in the Alola region dedicated to wildlife conservation. Faba attempts to summon Nihilego, which captures Lusamine, Lillie's mother, and takes her to its dimension. After Nihilego is defeated, he becomes much nicer, apologizing to Ash and his friends for his sinister ambitions ...
Lillie, Gladion, and Lusamine also make their way to the Crown Tundra to continue their search for Mohn. Lillie and her family eventually find Mohn at a cabin, but he has no memory of them due to having amnesia caused from the accident that sent him away from his family many years earlier.
After Lusamine uses Nebby's power to travel into the Ultra Wormhole to battle Necrozma, Lillie together with Selene summon the Legendary Pokémon Lunala in an effort to both restore Nebby's power and save Lillie's mother. After both Guzma and Lusamine fall out of the Ultra Wormhole, Necrozma emerges behind them.
The player then proceeds to battle through Aether Paradise with Hau and Gladion's help, eventually defeating Aether president Lusamine, who is revealed to be Gladion and Lillie's mother. Despite this, Lusamine and Guzma manage to open an Ultra Wormhole, with the former overclocking Nebby's powers, transporting them to Ultra Space, the Ultra ...
Back at the family mansion, Lusamine shows them Mohn's old office, where Lillie discovers her father's old Z-Ring and the inoperative Mythical Pokémon Magearna. Lillie goes to Kahuna Hala and asks for permission to use her father's Z-Ring, and they have a battle to see if Lillie can master Z-Moves.