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The Diptera is a very large and diverse order of mostly small to medium-sized insects. They have prominent compound eyes on a mobile head, and (at most) one pair of functional, membraneous wings, [1] which are attached to a complex mesothorax. The second pair of wings, on the metathorax, are reduced to halteres
English: Diagram of wing veins in males of pointed-winged flies (Diptera: Lonchopteridae) Italiano: Schema della nervatura alare nei maschi di Lonchoptera sp. (Diptera: Lonchopteridae) Date
The wings are moved by the rapid muscular contraction and expansion of the thorax. [11] The wings arise from the meso- and meta-thoracic segments and are similar in size in the basal groups. In more derived groups, the meso-thoracic wings are larger with more powerful musculature at their bases and more rigid vein structures on the costal edge. [7]
In the flexed wing, the outer squama of the alula is turned upside down above the inner squama, the latter not being affected by the movement of the wing. In many Diptera, a deep incision of the anal area of the wing membrane behind the single vannal vein sets off a proximal alar lobe distal to the outer squama of the alula. [38]
A single large wing from a species of Diptera in the Triassic (10 mm instead of usual 2–6 mm) was found in Australia (Mt. Crosby).This family Tilliardipteridae, despite the numerous 'tipuloid' features, should be included in Psychodomorpha sensu Hennig on account of loss of the convex distal 1A reaching wing margin and formation of the anal loop.
The majority of insects have two pairs of wings. Flies possess only one set of lift-generating wings and one set of halteres. The order name for flies, "Diptera", literally means "two wings", but there is another order of insect which has evolved flight with only two wings: strepsipterans, or stylops; [2] they are the only other organisms that possess two wings and two halteres. [6]