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In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". [3] [4] The term was introduced by Carl Nägeli in 1858. [5] [6] Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem.
In the stems of some Asterales dicots, there may be phloem located inwardly from the xylem as well. Between the xylem and phloem is a meristem called the vascular cambium. This tissue divides off cells that will become additional xylem and phloem. This growth increases the girth of the plant, rather than its length.
The combination of one xylem and one phloem strand adjacent to each other is known as a vascular bundle. [14] The evolution of vascular tissue in plants allowed them to evolve to larger sizes than non-vascular plants , which lack these specialized conducting tissues and are thereby restricted to relatively small sizes.
Plasmodesmata are also used by cells in phloem, and symplastic transport is used to regulate the sieve-tube cells by the companion cells. [citation needed] The size of molecules that can pass through plasmodesmata is determined by the size exclusion limit. This limit is highly variable and is subject to active modification. [5]
Xylem and Phloem. A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, engages in photosynthesis, stores nutrients, and produces new living tissue. [1]
There is also a tissue between xylem and phloem, which is the cambium. The xylem typically lies towards the axis ( adaxial ) with phloem positioned away from the axis ( abaxial ). In a stem or root this means that the xylem is closer to the centre of the stem or root while the phloem is closer to the exterior.
Phloem is a complex tissue, consisting of two main cell types, the sieve tubes and the intimately associated companion cells, together with parenchyma cells, phloem fibres and sclereids. [ 19 ] : 171 Sieve tubes are joined end-to-end with perforated end-plates between known as sieve plates , which allow transport of photosynthate between the ...
The pericycle is located between the endodermis and phloem in plant roots. In dicot stems, it is situated around the ring of vascular bundles in the stele. [1]