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The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers.They differ from mosses by being vascular, i.e., having specialized tissues that conduct water and nutrients, and in having life cycles in which the branched sporophyte is the dominant phase.
An interesting case is that of Psilotum, which has a (simple) protostele, and enations devoid of vascular tissue. Some species of Psilotum have a single vascular trace that terminates at the base of the enations. [2] Consequently, Psilotum was long thought to be a "living fossil" closely related to early land plants (rhyniophytes). However ...
The term "fern ally" included under Pteridophyta generally refers to vascular spore-bearing plants that are not ferns, including lycopods, horsetails, whisk ferns and water ferns (Marsileaceae, Salviniaceae and Ceratopteris). This is not a natural grouping but rather a convenient term for non-fern, and is also discouraged, as is eusporangiate ...
Some species have leaf-like structures called enations which have no vascular tissue except for a small bundle at the base. These are almost peg-like, stubby and are generally not considered true leaves, though they likely evolved from them. Members of Tmesipteris may appear to have leaves, but these are really phylloclades, or flattened stems.
More recent genetic studies have shown that the Lycopodiophyta are only distantly related to any other vascular plants, having radiated evolutionarily at the base of the vascular plant clade, while both the whisk ferns and horsetails are as much true ferns as are the Ophioglossoids and Marattiaceae. The Marattiaceae are a group of tropical ...
Psilotum is a genus of fern-like vascular plants.It is one of two genera in the family Psilotaceae commonly known as whisk ferns, the other being Tmesipteris.Plants in these two genera were once thought to be descended from the earliest surviving vascular plants, but more recent phylogenies place them as basal ferns, as a sister group to Ophioglossales.
Tracheids were the main conductive cells found in early vascular plants. In the first 140-150 million years of vascular plant evolution, tracheids were the only type of conductive cells found in fossils of plant xylem tissues. [5] Ancestral tracheids did not contribute significantly to structural support, as can be seen in extant ferns. [6]
Most non-vascular plants, as well as many lycophytes and most ferns, are homosporous (only one kind of spore is produced). Some lycophytes, such as the Selaginellaceae and Isoetaceae , [ 7 ] : 7 the extinct Lepidodendrales , [ 8 ] and ferns, such as the Marsileaceae and Salviniaceae are heterosporous (two kinds of spores are produced).