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A simple cladogram showing the evolutionary relationships between four species: A, B, C, and D. Here, Species A is the outgroup, and Species B, C, and D form the ingroup. In cladistics or phylogenetics, an outgroup [1] is a more distantly related group of organisms that serves as a reference group when determining the evolutionary relationships of the ingroup, the set of organisms under study ...
A nonoptimal cladogram will be selected if the program settles on a local minimum rather than the desired global minimum. [16] To help solve this problem, many cladogram algorithms use a simulated annealing approach to increase the likelihood that the selected cladogram is the optimal one. [17]
If the lineages in the phylogenetic tree stand for species, a polytomy shows the simultaneous speciation of three or more species. [6] In particular situations, they may be common, for example when a species that has rapidly expanded its range or is highly panmictic undergoes peripatric speciation in different regions.
Willi Hennig 1972 Peter Chalmers Mitchell in 1920 Robert John Tillyard. The original methods used in cladistic analysis and the school of taxonomy derived from the work of the German entomologist Willi Hennig, who referred to it as phylogenetic systematics (also the title of his 1966 book); but the terms "cladistics" and "clade" were popularized by other researchers.
Cladogram (a branching tree diagram) illustrating the relationships of organisms within groups of taxa known as clades. The vertical line stem at the base represents the last common ancestor . The blue and orange subgroups are clades, each defined by a common ancestor stem at the base of its respective subgroup branch .
For example, if we consider the crown-birds (i.e. all extant birds and the rest of the family tree back to their most recent common ancestor), extinct side branches like the dodo or great auk are still descended from the most recent common ancestor of all living birds, so fall within the bird crown group. [4]
Example: Saying that Tyrannosaurus rex had an eyeball is a level 1 inference because both extant members of the groups encompassing Tyrannosaurus rex have eyeballs, and eyeball sockets (orbital excavations) in the skull, the homology of which is well established, and the fossils of Tyrannosaurus rex skulls have similar morphology.
The green box (central) may represent an evolutionary grade (paraphyletic), a group united by conservative anatomical and physiological traits rather than phylogeny. The flanking red and blue boxes are clades (i.e., complete monophyletic subtrees). A grade is a taxon united by a level of morphological or physiological complexity.