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The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. [ 1 ] [ 2 ] [ 3 ] It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya . [ 4 ]
The two-empire system or superdomain system, proposed by Mayr (1998), with top-level groupings of Prokaryota (or Monera) and Eukaryota. [11] [12] The eocyte hypothesis, proposed by Lake et al. (1984), [13] which posits two domains, Bacteria and Archaea, with Eukaryota included as a subordinate clade branching from Archaea. [14] [13] [15]
Two kingdoms, Archaebacteria (archaea) and Eubacteria (for bacteria) were established. [22] Based on further studies, Woese, Otto Kandler and Mark Wheelis introduced the concept of "domain" in 1990 as the highest level of biological classification, and proposed the three-domain system consisting of Eucarya, Bacteria and Archaea. [23]
The word archaea comes from the Ancient Greek ἀρχαῖα, meaning "ancient things", [16] as the first representatives of the domain Archaea were methanogens and it was assumed that their metabolism reflected Earth's primitive atmosphere and the organisms' antiquity, but as new habitats were studied, more organisms were discovered.
The three-domain system adds a level of classification (the domains) "above" the kingdoms present in the previously used five- or six-kingdom systems.This classification system recognizes the fundamental divide between the two prokaryotic groups, insofar as Archaea appear to be more closely related to eukaryotes than they are to other prokaryotes – bacteria-like organisms with no cell nucleus.
Metagenomic analyses recover a two-domain system with the domains Archaea and Bacteria; in this view of the tree of life, Eukaryotes are derived from Archaea. [ 57 ] [ 58 ] [ 59 ] With the later gene pool of LUCA's descendants, sharing a common framework of the AT/GC rule and the standard twenty amino acids, horizontal gene transfer would have ...
In this scheme they reintroduced the classification with the division of prokaryotes superkingdom into two kingdoms, Bacteria (=Eubacteria) and Archaea (=Archaebacteria). This is based on the consensus in the Taxonomic Outline of Bacteria and Archaea (TOBA) and the Catalogue of Life .
Neomura (from Ancient Greek neo-"new", and Latin-murus "wall") is a proposed clade of life composed of the two domains Archaea and Eukaryota, coined by Thomas Cavalier-Smith in 2002. [1] Its name reflects the hypothesis that both archaea and eukaryotes evolved out of the domain Bacteria , and one of the major changes was the replacement of the ...