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In plants, ATP is synthesized in the thylakoid membrane of the chloroplast. The process is called photophosphorylation. The "machinery" is similar to that in mitochondria except that light energy is used to pump protons across a membrane to produce a proton-motive force. ATP synthase then ensues exactly as in oxidative phosphorylation. [28]
28.2% (sunlight energy collected by chlorophyll) → 68% is lost in conversion of ATP and NADPH to d-glucose, leaving; 9% (collected as sugar) → 35–40% of sugar is recycled/consumed by the leaf in dark and photo-respiration, leaving; 5.4% net leaf efficiency. Many plants lose much of the remaining energy on growing roots.
The Gibbs free energy is the energy available ("free") to do work. Any reaction that decreases the overall Gibbs free energy of a system will proceed spontaneously (given that the system is isobaric and also at constant temperature), although the reaction may proceed slowly if it is kinetically inhibited.
The efficiency of both light and dark reactions can be measured, but the relationship between the two can be complex. For example, the light reaction creates ATP and NADPH energy molecules, which C 3 plants can use for carbon fixation or photorespiration. [44] Electrons may also flow to other electron sinks.
To do this, it must release the absorbed energy. This can happen in various ways. The extra energy can be converted into molecular motion and lost as heat, or re-emitted by the electron as light (fluorescence). The energy, but not the electron itself, may be passed onto another molecule; this is called resonance energy transfer.
The Calvin cycle uses the chemical energy of ATP and the reducing power of NADPH from the light-dependent reactions to produce sugars for the plant to use. These substrates are used in a series of reduction-oxidation ( redox ) reactions to produce sugars in a step-wise process; there is no direct reaction that converts several molecules of CO 2 ...
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The ATP generated in this process is made by substrate-level phosphorylation, which does not require oxygen. Fermentation is less efficient at using the energy from glucose: only 2 ATP are produced per glucose, compared to the 38 ATP per glucose nominally produced by aerobic respiration. Glycolytic ATP, however, is produced more quickly.