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Anatomy of a eudicot leaf. The plant cuticle is a layer of lipid polymers impregnated with waxes that is present on the outer surfaces of the primary organs of all vascular land plants. It is also present in the sporophyte generation of hornworts, and in both sporophyte and gametophyte generations of mosses. [2]
The endodermis is the innermost layer of cortex in land plants. It is a cylinder of compact living cells, the radial walls of which are impregnated with hydrophobic substances (Casparian strip) to restrict apoplastic flow of water to the inside. [1] The endodermis is the boundary between the cortex and the stele.
The epidermis is the outermost cell layer of the primary plant body. In some older works the cells of the leaf epidermis have been regarded as specialized parenchyma cells, [1] but the established modern preference has long been to classify the epidermis as dermal tissue, [2] whereas parenchyma is classified as ground tissue. [3]
As a result, the Casparian strip is separated from the cytoplasm and the connection between the two ceases to be evident. In the third stage, a thick cellulose layer is deposited over the suberin, sometimes mainly on the inner tangential walls. The thickened wall, as well as the original wall in which the Casparian strip is located, may become ...
Many species of the genus Primula and ferns, such as Cheilanthes, Pityrogramma and Notholaena, as well as many genera of Crassulaceae succulent plants, produce a mealy, whitish to pale-yellow glandular secretion known as farina that is not an epicuticular wax, but consists largely of crystals of a different class of polyphenolic compounds known as flavonoids. [5]
It is an insoluble substance with waterproof quality. Cutin also harbors cuticular waxes, which assist in cuticle structure. [ 1 ] Cutan, the other major cuticle polymer, is much more readily preserved in fossil records. [ 2 ]
The cells in this layer are dead and abundant in suberin, preventing water loss from the tissues below. Suberin can also be found in various other plant structures. For example, they are present in the lenticels on the stems of many plants and the net structure in the rind of a netted melon is composed of suberised cells.
As the leaf grows, the pavement cells will also grow, divide, and synthesize new vacuoles, plasma membrane parts, and cell wall components. A thick external cell wall influences the direction of growth by impeding expansion towards the outside of the cell and instead promote expansion parallel to the epidermis layer. [7]