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The two-domain system is a biological classification by which all organisms in the tree of life are classified into two domains, Bacteria and Archaea. [ 1 ] [ 2 ] [ 3 ] It emerged from development of knowledge of archaea diversity and challenges the widely accepted three-domain system that classifies life into Bacteria, Archaea, and Eukarya . [ 4 ]
Metagenomic analyses recover a two-domain system with the domains Archaea and Bacteria; in this view of the tree of life, Eukaryotes are derived from Archaea. [ 58 ] [ 59 ] [ 60 ] With the later gene pool of LUCA's descendants, sharing a common framework of the AT/GC rule and the standard twenty amino acids, horizontal gene transfer would have ...
Two kingdoms, Archaebacteria (archaea) and Eubacteria (for bacteria) were established. [22] Based on further studies, Woese, Otto Kandler and Mark Wheelis introduced the concept of "domain" in 1990 as the highest level of biological classification, and proposed the three-domain system consisting of Eucarya, Bacteria and Archaea. [23]
According to the domain system, the tree of life consists of either three domains, Archaea, Bacteria, and Eukarya, [1] or two domains, Archaea and Bacteria, with Eukarya included in Archaea. [3] [4] In the three-domain model, the first two are prokaryotes, single-celled microorganisms without a membrane-bound nucleus.
In 1990 Carl Woese et al. introduced the three-domain system. [35] [36] The prokaryotes were split into two domains, the archaea and the bacteria, while the eukaryotes become a domain in their own right. The key difference from earlier classifications is the splitting of archaea from bacteria.
Archaea were initially classified as bacteria, receiving the name archaebacteria (/ ˌ ɑːr k i b æ k ˈ t ɪər i ə /, in the Archaebacteria kingdom), but this term has fallen out of use. [5] Archaeal cells have unique properties separating them from Bacteria and Eukaryota. Archaea are further divided into multiple recognized phyla.
The two-empire system would later be expanded to the three-domain system of Archaea, Bacteria, ... the diagram below is an 'organization chart', ...
The point of fusion (marked "?") below LECA is the FECA, the first eukaryotic common ancestor, some 2.2 billion years ago. Much earlier, some 4 billion years ago, the LUCA gave rise to the two domains of prokaryotes, the bacteria and the archaea. After the LECA, some 2 billion years ago, the eukaryotes diversified into a crown group, which gave ...