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Overlying the boney elements are a series of scutes, which are made of keratin and are a lot like horn or nail tissue. In the center of the carapace are five vertebral scutes and out from these are four pairs of costal scutes. Around the edge of the shell are 12 pairs of marginal scutes.
Dermal scutes are also found in the feet of birds and tails of some mammals, and are believed to be the primitive form of dermal armour in reptiles. The term is also used to describe the heavy armour of the armadillo and the extinct Glyptodon , and is occasionally used as an alternative to scales in describing snakes or certain fishes, such as ...
Scutes on a crocodile. Reptile skin is covered with scutes or scales which, along with many other characteristics, distinguish reptiles from animals of other classes. They are made of alpha and beta-keratin and are formed from the epidermis (contrary to fish, in which the scales are formed from the dermis).
The "central scutes" extend over the dorsal mid line of the carapace from head to tail, with the "costal scutes" running along each side of the central scutes. The "marginal scutes" run along the outer sides of the shell, and the "nuchal scutes" are found in the area directly behind the turtle's head.
This species can be recognized by its black or brown oval carapace (upper shell) with distinctive low keels on the second and third scutes. Yellow spots on the side of its head give this species its common name. These spots are most prominent in juveniles and fade with age. Females can be up to twice the size of males.
The scales and scutes of birds were thought to be homologous to those of reptiles, [4] but are now agreed to have evolved independently, being degenerate feathers. [ 5 ] [ 6 ] Carcharodontosaurid theropod dinosaur Concavenator , is known to have possessed these feather-derived tarsal scutes.