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Secondary xylem is formed during secondary growth from vascular cambium. Although secondary xylem is also found in members of the gymnosperm groups Gnetophyta and Ginkgophyta and to a lesser extent in members of the Cycadophyta , the two main groups in which secondary xylem can be found are:
Stained Xylem Vessel in Red/Orange (x400 magnification) In secondary xylem – the xylem that is produced as a stem thickens rather than when it first appears – a vessel element originates from the vascular cambium. A long cell, oriented along the axis of the stem, called a "fusiform initial", divides along its length forming new vessel elements.
The vascular cambium produces secondary xylem on the inside of the ring, and secondary phloem on the outside, pushing the primary xylem and phloem apart. The vascular cambium usually consists of two types of cells: Fusiform initials (tall, axially oriented) Ray initials (smaller and round to angular in shape)
Plant cells will increase the production of lignin generating enzymes when stressed by some pathogens, further lignifying the secondary cell wall. Increased lignin content is particularly effective at resisting vascular pathogens that use the secondary xylem to spread.
Between the xylem and phloem is a meristem called the vascular cambium. This tissue divides off cells that will become additional xylem and phloem. This growth increases the girth of the plant, rather than its length. As long as the vascular cambium continues to produce new cells, the plant will continue to grow more stout.
Secondary growth thickens the stem and roots, typically making them woody.Obstructions such as this metal post and stubs of limbs can be engulfed. In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems ...
A cambium (pl.: cambiums or cambia), in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from which phloem, xylem, or cork grows by division, resulting (in woody plants) in secondary thickening.
The cambium tissues, i.e., the cork cambium and the vascular cambium, are the only parts of a woody stem where cell division occurs; undifferentiated cells in the vascular cambium divide rapidly to produce secondary xylem to the inside and secondary phloem to the outside.